Principal Investigator

Dan Speiser
Dan is a graduate of East Lansing High School ('99) and Carleton College ('03). He earned a PhD as a member of Sonke Johnsen's Sensory Ecology lab at Duke University ('10) and was a post-doc in Todd Oakley's Evolutionary Biology lab at UC Santa Barbara (2011-14). Dan started as an Assistant Professor at the University of South Carolina in 2014, was promoted to Associate Professor in 2020, and then promoted to Professor in 2026. He's not sure when  mollusks became his muse, but it probably dates back to childhood battles with marauding garden slugs or a magical encounter with an Aplysia in an undergraduate invertebrate biology course held in the dead of a Minnesota winter.

Graduate Students

Maddy Janakis
Maddy graduated in 2020 from DePaul University with a BS in Biology and Environmental Science. She is currently a PhD candidate busy researching how crab vision is suited to dynamic light conditions using color and polarized signals. She's also interested in crab behavior in different contexts. She loves her cat, Frankie, her turtle, Al Capone, and, of course, her extended family of crabs. Maddy enjoys Glee, ghosts, Kesha, beach days and driving her fellow lab members to the brink of insanity. Find her on twitter @JanakisMaddy and on Bluesky @maddy-janakis.bsky.social
Adriana Halvonik-Sanchez
Adriana graduated from the University of Costa Rica in 2022 with a BS in Biology. She is currently working on her PhD at USC where she is studying the ecological niches and evolutionary history of a group of closely related chitons that coexist throughout the Americas. She is also developing a bilingual field guide for the taxonomic identification of chitons in Central America. In her free time, she enjoys cooking, exploring tidepools in rocky beaches, and spending time with her cats, Toe Beans and Andrew, and her dog, Camo.
Quaid Pendleberry
Quaid graduated in 2020 with a BS in Biology from Marshall University.  Quaid’s undergraduate work focused on big river ecology and salamander population dynamics. Since then, they have been working as a field biologist on a variety of projects, mainly pertaining to migratory birds.  After working for several years, they are excited to restart their academic life as a graduate student focusing on behavior and visual ecology.  Quaid enjoys spending time with their partner and two cats, Ebi and Mochi. Quaid also enjoys crawling around in the woods, scuba diving, and taking pictures of any critter they can.
Haley Hines
Haley graduated from Virginia Tech in 2025 with a BS in Biological Sciences and a minor in Professional Sales. Her undergraduate research involved using bioinformatics to study the evolution of sodium channels in echinoderms. She is now a PhD student at USC studying functional neurobiology in scallops. She is also interested in distributed vision across various taxa. Haley enjoys reading fantasy books, cheering on her favorite sports teams, spending time outside, and feeding her lizard, Phoebe.

Undergraduate Researchers

C. Wynn Ravan
Wynn Ravan is a sophomore Marine Science major with a concentration in biological oceanography. He is looking to gain experience relevant to a career in coral conservation through DNA analysis and phylogenetics. He is currently planning on pursuing a graduate degree in Marine Science. Outside of school, he has a newfound passion for SCUBA diving and enjoys spending time playing with his cat Calypso, cooking, and being on/near the water.